What does a crumbling cork say about storage conditions?

The following appeared in a recent post:

"the cork on the 1990 dunn crumbled when he tried to pull it out, so storage was most likely the issue."

What does storage have to do with cork condition? I mean, aside from the point that if the cork does not remain wet, it will likely shrink and allow air to enter the bottle.

I would think that crumbling or not crumbling depends mostly on the quality of the cork selected by the bottler.

I searched this topic and could not find any previous answer to the question. Thanks in advance.
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